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Urban environments are also attractive due to the large prey concentrations and lower amounts of predation. Nephila produce large asymmetric orb webs up to one and a half meters in diameter.
The hub of the web is not positioned centrally but rather in the upper section; this means that the majority of the sticky capture strands are found in the lower section of the web.
There is a barrier web structure on either side of the main web to mitigate this risk. Yellow threads of their web shine like gold in sunlight.
Carotenoids are the main contributors to this yellow color, but xanthurenic acid , two quinones and an unknown compound may also aid in the color.
The spider is able to adjust pigment intensity relative to background light levels and color; the range of spectral reflectance is specifically adapted to insect vision.
The webs of most Nephila spiders are complex, with a fine-meshed orb suspended in a maze of non-sticky barrier webs. As with many weavers of sticky spirals, the orb is renewed regularly if not daily, apparently because the stickiness of the orb declines with age.
When weather is good and no rain has damaged the orb web , subadult and adult Nephila often rebuild only a portion of the web. The spider will remove and consume the portion to be replaced, build new radial elements, then spin the new spirals.
This partial orb renewal is distinct from other orb-weaving spiders that usually replace the entire orb web.
In it was discovered that the web of Nephila antipodiana contains ant-repellent chemicals to protect the web. Typically, the golden orb-weaver first weaves a non-sticky spiral  with space for two to twenty more spirals in between the density of sticky spiral strands decreases with increasing spider size.
When she has completed the coarse weaving, she returns and fills in the gaps. Whereas most orb-weaving spiders remove the non-sticky spiral when spinning the sticky spiral, Nephila leave it.
This produces a " manuscript paper " effect when the orb is seen in the sun: The circular-orb portion of a mature N. In relation to the ground, the webs of adults may be woven anywhere from eye-level upwards high into the tree canopy.
The orb web is usually truncated by a top horizontal support strand, giving it an incomplete look. Adjacent to one face of the main orb there may be a rather extensive and haphazard-looking network of guard-strands suspended a few inches distant across a free-space.
This network is often decorated with a lumpy string or two of plant detritus and insect carcasses clumped with silk.
At least one reference explains the suspended debris-chain as a cue for birds to avoid blundering into and destroying the web.
The golden silk orb weaver targets many different organisms as prey, ranging from small flies and beetles to larger cicadas and locusts. As a result of their strong web structure small birds and bats can also become trapped and fed upon.
Whilst most of the captured prey is relatively small compared to Nephila so the majority of biomass consumed comes from larger more rare prey.
Throughout the maturation process spiders exhibit different feeding behaviours relative to their web type. Originally after birth the spiderlings inhabit a communal web and eat dead siblings and dead prey but do not attack any live organisms.
After they move to individual webs the spiderlings begin to attack small live prey. As they increase in size they become more ambitious in their prey capture attempts through larger and stronger webs and greater attack confidence.
Spiders are notified that potential prey has been caught in the web through vibrations along strands, these can be followed to the prey location on the web and be used to estimate prey size.
Small prey like flies are captured alive then removed from the web and wrapped in silk. This occurs with smaller prey because they do not present a threat to the spider when alive.
Prey that is seen as potentially threatening will be bitten to be subdued before wrapping. Medium-sized prey will receive bites that are sustained until prey is immobilised and then be covered in silk.
Finally, the largest captured prey receive the —bite and back off— treatment where a single lunging bite is made before the spider withdraws and waits for the venom to take effect before wrapping.
This lunging bite is often repeated to speed up the process of immobilisation. Nephila also create caches of food for storage, these can be found above the hub of the web and contain up to 15 prey items.
These items are arranged in a line vertically and are wrapped in silk to reduce dehydration. Caches are created and grow when prey is readily available and there is more biomass available for consumption than is required by the spider.
The purpose of caches is to have a backup food source when prey is scarce and occasionally to provide bait to attract more prey to the web.
Nephila display large sexual size dimorphism with females being greatly larger than males, sometimes up to ten times the size.
Smaller males are quicker and more nimble, allowing them to be able to catch the females more easily as well as escape when threatened.
Larger males may have to wait for the female to come close due to their slower speed. Larger individuals reduce the success of these plugs, allowing for multiple mating and reducing the risk of genital mutilation.
When males are fully mature they leave their webs to search for a suitable female, often using web characteristics to identify potential mates.
Females are most likely to mate directly after their final molt at the beginning of adulthood and later in adulthood when they are feeding.
When males approach females they are often feeding, allowing the males to get closer without an aggressive response and also meaning the female is not moving.
On approach the male makes himself known by tapping on a web strand to ensure the female is amenable before proceeding to mate.
When met with aggression males will stop approaching and remain in the same location until the female relaxes or they retreat.
Females engage in multiple mating but there is no benefit to the offspring as a result of this; however the energy cost of repelling a male is higher than that of allowing him to copulate.
Sexual cannibalism is uncommon in Nephila as a result of male mating behaviours. By copulating when females are immobile after molting or inactive due to feeding, the males increase their chances of survival.
Males also approach from the side of the web opposite the female, increasing the odds of a successful approach. Female Nephila produce an egg-sac in the surrounding environs of the web to protect their eggs.
The eggs are deposited on a silk platform then are covered in loose silk to form a sac. This sac is firmly attached to surrounding vegetation so that it is hidden from the view of predators.
These sacs can contain from to eggs, depending on mating success and particular species. Nephila change their body positioning relative to the sun to maintain internal temperatures at an optimal level.
As ambient temperatures increase the spider will position itself so the abdomen shades the cephalothorax from the sun. Spiders may also hang from their hind legs as a result of the heat due to a loss of hydrostatic pressure.
Conversely as temperatures cool down the spider will position itself perpendicular to the sun to retain as much heat energy as possible. Predation of Nephila is relatively uncommon, when it does occur the main group affected are the juvenile individuals.
The major predators are birds but wasps and damselflies also prey upon smaller juveniles. To combat this females aggregate their webs to form a large complex, discouraging aerial predators for fear of entanglement.
The frequent rebuilding or abandoning of webs by Nephila may be a tactic for controlling Argyrodes. Spiny orb-weaver spiders of the genus Gasteracantha also inhabit the webs of Nephila as kleptoparasites, too.
Egg-sacs generally remain free from both predation and parasitism, often due to the close proximity of the mother and how well it is hidden.
Moth larvae of genus Anatrachyntis have been known to prey on Nephila eggs but this is the only well known and researched case. Nephila spiderlings leave the egg-sac as a result of environmental cues, often warmer and wetter conditions in spring.
They then live on a communal web eating dead siblings and web debris for around a week before dispersing to make individual webs. Young spiders do not generally build yellow-colored silk, and the young Nephila themselves can be easily mistaken for young Orchard Spiders Leucauge in general color and shape both genera sport silver stripes or patches on their abdomens, described in some references as a form of heat control.
The best distinction between Leucauge and Nephila juveniles is web structure: Leucauge tends to build a horizontal orb that is a perfect circle, whereas Nephila build vertical, elliptical orbs that are incomplete missing the portion of the orb over the hub, the center where the spider sits.
Nephila seem to prefer more open habitat such as second-growth scrub or forest edges. Fences or building overhangs often do just as nicely.
In addition, young spiders demonstrate vibrational motion when approached by a predator. If a predator persists in an attack, the spider will either run to a web-support strand and thus to nearby vegetation, bail out of the web on a silk line that remains connected to the web, or jump from the web after inducing oscillations in the web that aid the jump.
Once they are juvenile, Nephila inhabit their individual webs then begin the growth by molting process. Easy to use, no fuss and quality. This also goes for the games that are offered for playing on mobile devices.
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